The new partners will each contribute £40 million in the UK Centre for Medical Research and Innovation, bringing the total committed funding for the planned London institute to £700 million ($1.14 billion).
UKCMRI has said genomics and other technologies will be applied toward its four key goals: research excellence, supporting the UK's biomedical research effort, recruiting and training researchers, and fostering translation of discoveries into new treatments against disease.
The application is expected to be heard before the end of the year by members of the Camden Council's Development Control Committee, following a formal consultation by the council. If approved, construction will begin next year, with completion set for 2015.
The UK Centre for Medical Research and Innovation will maintain "a strong focus" on cancer, heart disease, and stroke as well as disorders of the immune system and later-life diseases of the nervous system, partners in the planned £600 million institute said.
The coalition headed by David Cameron's Conservative Party said that it remains committed to the project, promising to spend the remaining £233 million set aside for UKCMRI by the UK government in phases over the next five years.