In March 1937, Hans Krebs and his colleague conducted an experiment observing the metabolic rate decline of the minced breast of a freshly killed pigeon suspended in solution. According to an article in The Scientist, by adding a salt of citric acid, they found they could keep the tissue "alive" for three times as long, and additional experiments showed the cyclical nature of the pathway which regenerates citric acid with each cycle and releases ATP.

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